of research This study has received assistance from National Natural Science Foundation of China (No. 32172932, 32002346), Key Program of Natural Science Foundation of Heilongjiang Province of China (No. ZD2021C003), China Agriculture Research System of MOF and MARA (No. CARS-35), Distinguished Professor of Longjiang Scholars Support Project (No. T201908) and Heilongjiang Touyan Innovation Team Program. CRediT authorship contribution statement Xue-Yan Dai: Conceptualization, Methodology, Investigation, Formal analysis, Writing – original draft, Writing – review & editing. Jia Lin: Formal analysis, Writing – review & editing. Shi-Yong Zhu: Formal analysis, Writing – review & editing. Jian-Ying Guo: Formal analysis, Writing – review & editing. Jia-Gen Cui: Writing – review & editing. Jin-Long Li: Conceptualization, acquisition, Supervision, Writing – review & editing.