Funding: This research was jointly supported by National Key R&D Program of China (2016YFD0501210, 2017YFD0500505), the Program of National Agricultural Research Outstanding Talents of China (2015), Hubei Provincial Technology and Innovation Program (2016ABA121, 2017AHB062), Natural Science Foundation of Hubei Province (2016CFA070), the Hubei Hundred Talent Program, and Texas AgriLife Research (H-8200).