Conceptualization, Q.L. and T.Z.; methodology, N.Z.; software, X.Z. and X.D.; validation, R.H., H.X. (Hongsen Xu), F.C. and H.X. (Huili Xue); formal analysis, Q.L. and N.Z.; investigation, Q.L., N.Z. and T.Z.; resources, X.D.; data curation, Q.L. and F.Z.; writing—original draft preparation, Q.L.; writing—review and editing, N.Z. and T.Z.; visualization, R.H., H.X. (Hongsen Xu), F.C., H.X. (Huili Xue) and F.Z.; supervision, X.D. and T.Z.; project administration, Q.L., N.Z. and T.Z.; funding acquisition, X.D. and T.Z. All authors have read and agreed to the published version of the manuscript. This research was funded by key agricultural projects of major science and technology programs of the Science Technology Department of Zhejiang Province [grant no. 2015C02028]; key scientific and technological grant of Zhejiang for breeding new agricultural varieties [grant no. 2021C02069-8]; key research and development project on special agricultural new variety breeding of major science and technology program of the Zhejiang Province [grant no. 2016C02055-4].