Drs Tao Wu, Kang Li and Yang Lyu contributed equally to this work and should be considered co-first authors. This work was jointly supported by the National Key R&D Program (grant number 2017YFD0500505, 2016YFD0501210), the Program of National Agricultural Research Outstanding Talents of China (2015), Hubei Provincial Key R&D Program (2019ABA083), the Open Project of Hubei Key Laboratory of Animal Nutrition and Feed Science (grant number 201805) and Texas A&M AgriLife Research (H-8200). We thank our students and technicians for their contributions to this research.
