Conceptualization, S.G. and B.D.; methodology, L.H., Y.Z. and J.N.; software, S.G. and L.H.; validation, Z.Z., and P.L.; formal analysis, S.G., Z.Z. and B.D.; investigation, S.G., L.H., Y.Z. and J.N.; resources, Z.Z.; data curation, S.G., C.L. and Z.Z.; writing—original draft preparation, S.G. and L.H.; writing—review and editing, C.L., Z.Z., P.L. and B.D.; visualization, Y.Z. and J.N.; supervision, B.D.; project administration, S.G. and B.D.; funding acquisition, S.G. and B.D. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Natural Science Foundation of China (No. 32172757), the National Key R&D Program of China (No. 2016YFD0501202), the National Science Foundation of Hubei Province (2021CFB559) and the Open Project of Hubei Key Laboratory of Animal Nutrition and Feed Science (No. 201902).