This research was funded by the National Major Projects of China (2016ZX08010004-004), the National Natural Science Foundation of China (31771418, 32272126), the Natural Science Foundation of Hubei China (2016CFB549), the Fundamental Research Funds for Central Universities, HUST (2021XXJS070, 3004170157) and Wuhan Knowledge Innovation Project (2022020801010073). This work was also supported by the Opening Fund of Key Laboratory of Forage and Endemic Crop Biology, Ministry of Education (FECBOF2021002), the Opening Fund of Hubei Key Laboratory of Bioinorganic Chemistry & Materia Medica (BCMM202205), the Start-Up Research Funding of Wuhan Polytechnic University (2021RZ100, 53210052172) and the Research Project of Wuhan Polytechnic University (2023Y24).