Conceptualization, D.X., A.Y. and S.C. (Shuiyuan Cheng); methodology, X.Z. and S.C. (Shaoyu Chen); investigation, X.T. and T.Q.; formal analysis, Y.X., X.T. and T.Q.; writing—original draft preparation, Y.X.; writing—review and editing, M.W. and D.X.; funding acquisition, X.C. and Y.X. All authors have read and agreed to the published version of the manuscript. This work was supported by grants from Hubei Key Laboratory of Food Crop Germplasm and Genetic Improvement (2020lzjj05); School of Modern Industry for Selenium Science and Engineering, Wuhan Polytechnic University (Se1-202111) and Scientific Research Program of Education Department of Hubei Province (Q20221613).